nominate Lesser Black-backed Gull (L. f. fuscus)

(last update: 26-11-2014 )

Coordinators:
Amir Ben Dov (Israel)
Hannu Koskinen (Finland)
Mars Muusse (the Netherlands)

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adult fuscus: March


Gulls at Ashdod Israel, 16 March 2010.

In northern Europe, several ringing programmes were started to get more information about the feeding aspects, migration and behaviour of Larus fuscus fuscus. One of such ringing programmes is executed in the three northern counties of Norway: Nordland, Troms and Finnmark.
Morten Helberg, Geir Systad, Ingve Birkeland, Nils Lorentzen & Jan Bustnes published an article about this research in Ardea 97, 2009, titled: Migration patterns of adult and juvenile Lesser Black-backed Gulls Larus fuscus from northern Norway. The complete PDF can be found HERE. The summary for juveniles can be found in the fuscus 1cy October section, the summary for adults can be found in the fuscus adult October section.

Nordland - Hortavær

Hortavær, in the very south of the county Nordland, had a homogenous population of about 400 pairs of fuscus type birds in 2005 (birds showing blackish upperparts). In this population, only one or two paler birds have been observed; one of these paler birds may possibly have raised chicks in a mixed pair with fuscus. In 2008 a new population of 200 pairs was discovered nearby Hortavær; and also on this location was one pale non-breeding bird.

Table 1: results of the population study of northern Lesser Black-backed Gull (Larus fuscus fuscus) in the Hortavær Archipelago 2005-2008.
2005
2006
2007
2008
total
# nests
378
385
135
288
-
# colour ringed adults
83
42
13
16
154
# ringed pulli
371
226
52
94
743
# colour ringed pulli
246
218
46
94
604
# young late July
400-500
350-400
150
150-180
1050-1180
# returned birds from 2005
-
0
9
25
34
# returned birds from 2006
-
-
0
2
2

Between 2005-2007, 509 birds have been ringed, and 26 birds were rediscovered. Despite being a colony of pure fuscus birds, several individuals were found following the western migration route, through the North Sea, south along the Atlantic coast of the Iberian peninsula and north Africa. Here are two example males from this population: J309 and J130. J309 (thumbnail left, click to enlarge), is a perfect illustration of jet-black adult nominate fuscus, breeding on Hortavær, Nordland. This Archipelago contains a pure population of such dark fuscus birds. Complete moult still not started by mid-July. J130 (thumbnail right), illustrates type of breeding habitat for this colony.

And also two example females from this population: J303 (left) and J127 (right). Again these birds illustrate nicely the darkness of the upperparts, leaving almost no contrast between the tone of the wing-coverts and the medial black bands on the primaries. By clicking the thumbnail of J127, you also find a map with migration strategy followed by this bird, which was seen in western Europe and Morocco. More details about the upperpart grey tone for various Lesser Black-backed Gull colonies throughout Europe, you can look at the fuscus adult August section. Here, you will see the findings of E Barth, Norwegian scientist who plotted grey tones in the 1960-1970's. Much of what he wrote, still is true in Norway.

In fuscus colonies, like in other white-headed gull species, you can find 3cy birds which return to inspect their future breeding grounds and feeding grounds, and some may even try to participate in courtship. In Hortavær, several 3cy birds, ringed in 2005 and 2006 have been seen back in the colony. These thumbnails give some illustrations. Probably, most of such 3cy birds involve males. Above left bird is J304, caught on the nest as 3cy fuscus. The right thumbnail shows 3cy J054, ringed as pullus in 2005 and returning in the colony in 2007. Below left is 3cy J229 and below right is 3cy J214, of which is included a map with migration strategy followed by this bird (seen in N Germany and Belgium).
Harry Vercruijsse (1999), who did research in the Netherlands on Herring Gulls in the dunes, found the breeding attempts by such young birds not really successful in their first year back in the colony. Birds in 3cy may occupy a territory, but real breeding attempts often start the next year. Still, it may be a good strategy to return at such early age, to have advantage over other males regarding best breeding locations and feeding grounds. Similar results were found by Chabrzyk & Coulson (1976) on the Isle of May, SE Scotland.
[HJP Vercruijsse, Zilvermeeuwen uit de duinen van Schouwen, 1999. Publication ibn dlo, Tilburg, ISBN 9090131671.
G Chabrzyk & JC Coulson, Survival and recruitment in the Herring Gull Larus argentatus, 1976. J of An Eco 45: 187-203.] (thumbnail left: 3cy J243).

Pale-mantled adults in Hortavær, Nordland. This Archipelago contains a pure population of dark fuscus birds, but sometimes a pale-mantled bird is seen here as well. Maybe there has been one breeding attempt of mixed fuscus type x pale-mantled bird.

Troms & Finnmark

North of the county Nordland lies Troms and the northernmost county is Finnmark. Starting in the south, Svartskjeran near Ibestad is the first colonies that was visited to measure and colour-ring Lesser Black-backed Gulls. Table 2 gives the breeding population for the counties Troms and Finnmark.

Table 2: results of the population study of northern Lesser Black-backed Gull (Larus fuscus fuscus) in Troms and Finnmark 2005-2008.
.
Max pairs
Ringed pulli
.
.
2005
2006
2007
2008
Svartskjeran
10 (fuscus)
13
10
3
2
Lemmingvaer
45 (mixed)
70
89
71
4
Froholman
40 (mixed)
47
67
46
2
Eggloysa
8 (mixed)
9
2
10
3
Kvannholmen
20 (mixed)
14
13
37
2
Musvaer / Auvaer
25 (mixed)
na
na
32
9
Sandvaer
6 (mixed)
0
6
4
2
Nordfugloy
100 (mixed)
8
57
61
25
Matmorstua - Loppa
10 (mixed)
7
12
5
0
Svarthellaren - Loppa
30 (mixed)
13
47
33
0
Total:
294
181
303
302
49

Click on Troms-Finnmark map below for larger image.

Troms - Svartskjeran / Ibestad

This small colony with maximum 10 breeding pairs consists of pure fuscus birds. All colonies north of Svartskjeran hold mixed populations with both fuscus type birds and pale-mantled birds. The island Svartskjeran was visited from 2005-2008 and 28 birds have been ringed here. The thumbnails of J000, J0R1 and J0R2 represent the type of birds breeding here, with jet-black upperparts. One of these example birds is J0R1, a bird ringed on the nest in 2005. This individual has been recorded several times in February and March 2008 at Ashdod, Israel. J0R1 follows the eastern migration route, earlier described by Kilpi and Saurola in 1984 for nominate fuscus.
Single observations of colour ringed birds from N Norway, e.g. observations from central inland Norway, the Baltic Sea and Israel demonstrate the eastern migration route (eastern flyway through Finland, Black Sea, Israel to the Rift Valley into the African Great Lakes). However, there were also observations of Norwegian birds from 'pure fuscus colonies' in England, which later that winter turned up in Morocco and another 'pure fuscus' bird in Portugal.

Troms - Lemmingsvær

Lemmingsvær holds one of the larger colonies in northern Norway, and ringing started already in 2002, with now 226 birds ringed in the period 2002-2008. Maximum breeding pairs is 45 birds for this location, with about 50% of these breeders being pale-mantled birds (origin either from graellsii populations from the west or intermedius birds colonising from southern Norway).
Example birds are JM82, JM85 (female) and JM88 (male). JM82 was not sexed. The complete moult has still not started by mid-June in this bird. It is obviously a pale-mantled bird, with a migration strategy much in line with what can be expected for intermedius birds from southern Norway or graellsii birds from the U.K. You can view the map by clicking the thumbnail of JM82.

Troms - Froholman / Eldhusholmen

The ringing programme at Froholman started in 2005 and within three years (2005-2007) 8 adults were caught on the nest. The catches of juveniles were better, with 162 juveniles colour ringed in the period 2005-2008; although last year only 2 birds were caught. 2008 has been a dramatic year regarding juveniles in all north Norwegian populations. None of the adults have been rediscovered outside the breeding range, up to May 2008, but three juveniles were seen again.

One of these birds is shown here as well, J916. This bird was seen in Israel and one year later in Egypt, nicely demonstrating the migration route already known for nominate fuscus from Finland. When J916 turned up in Egypt in February 2008 as 3cy bird, the upper-part grey tone was very dark, supporting the idea this is an offspring of pure fuscus.

As the first thumbnail shows, the population on Froholman obviously holds both pale-mantled birds and very dark fuscus-type birds.

Troms - Nordfugløy

The most northern island in the county Troms is Nordfugløy. Here you find a Lesser Black-backed Gull colony in cloudberry mires, about 300 meters above sea level. The oldest data about the presence of this colony are from 1963 when 125 pairs were estimated. Still, there was nothing known concerning the subspecies composition before the first adult birds were caught and colour-ringed in 2002. The breeding population varies a lot in numbers between the years, with a minimum of 15-20 pairs in 2002, and with a maximum of 100 pairs in 2006 and 2007. Nowadays, with about 100 pairs, it is the largest colony of Lesser Black-backed Gulls in northern Norway.

The island also hosts a breeding population of 100+ pairs of arctic skuas and 200+ pairs of Common Gulls. There is only one island in this ringing program which is higher up north: Loppa Island, which is a similar colony at about 200 meters above sea level.

1cy fuscus in August, ringed in Finland. (81365 bytes)The ringing program on Nordfugløy started in 2000, when juveniles were ringed. From 2000 to 2008, 172 juveniles were ringed here. Up to May 2008, 20 of these juveniles were seen outside the breeding range. Between 2002 and 2007, 17 adults were ringed as well, with 7 rediscoveries. This is a remarkable high recovery rate, certainly due to the predominant western migration route chosen by the adults breeding on this island. Pale-mantled birds from Nordfugløy have been rediscovered in Portugal, Spain, Italy and Libya. One dark fuscus-type bird has been observed frequently in Israel, J0X0 (see thumbnail right above).

adult March - description

By March many birds migrate north from nthe winter areas in Africa and pass Israel, where all images have been taken on this webpage. March is by far the best month to score ringed fuscus in Israel. In the January section and in the February section you can see that in ringed adult fuscus (provenance on age and location), the last phase of the complete moult includes growing P10. March and April are months used for migration and since fuscus is a long distance migrant one may expect no moult in this period. This appears to be true regarding the flight feathers. The list shows the primary moult scores for ringed birds and unringed birds of which the full set of primaries is visible in images.

"Primary moult scores for ringed and unringed fuscus in March"

P10 fully grown; P1-P10 present. PMS=0.
white ring: C1N9, C3Y1, C5P4, C5P8, C9XX, CCJ9, CE41;
red ring: C7EP, CE43, A170;
blue ring: J0X0;
black ring: J449;
ringed birds: 12.
unringed birds: 17.

It is difficult to determine exact moult stages in the wing-coverts in birds that are as dark as adult fuscus. For March, with the complete moult just finished, it may sound logic not to expect to much moult in the wing-coverts. And this is true for Finnish fuscus as far as we have checked the birds on this site. But interestingly, two birds, both from far northern Norway showed wing-covert moult: J0X0 and J20E.


March 13 2014. "the gathering".


Last days of March or sometimes the first week of April: Larus fuscus circling in groups to migrate north to the breeding grounds.


Spring Baltic Gulls in Israel. Picture: Amir Ben Dov.


Larus fuscus fuscus, March 21 2009, Yavne 4 water storage ponds, Ashdod, Israel. Picture: Amir Ben Dov.


Larus fuscus fuscus, March 21 2009, Yavne 4 water storage ponds, Ashdod, Israel. Picture: Amir Ben Dov.


Larus fuscus fuscus, March 21 2009, garbage transfer point area in Ashdod, Israel. Picture: Amir Ben Dov.

1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult C06Z March 08 2011, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult March 12 2011, Ashdod, Israel. Picture: Amir Ben Dov. Courtship.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult C09W March 29 2013, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult March 12 2011, Ashdod, Israel. Picture: Amir Ben Dov. Courtship.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus ad C0K5 March 16 2010, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 28 2013, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus 11-15+CY C0PE March 08 2011, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 02 2012, Ashdod, Israel. Picture: Amir Ben Dov. P2 new, P3 growing. Active moult in coverts.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus ad C172 March 17 2010, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 28 2013, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus 16CY C1EY March 29 20143, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 02 2012, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult C1M2 March 2013 & 2014, Ashdod, Israel. Picture: Amir Ben Dov. Complete moult finished. L. f. fuscus adult March 18 2011, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult C1N9 March 21 2009, Ashdod, Israel. Picture: Amir Ben Dov.Complete moult finished. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 02 2012, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult C2SN March 12 2011, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 23 2012, Eilat, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult C2VU March 21 2009, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult March 25 2011, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus ad C37Z March 02 2012, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult March 25 2011, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult C3.59 March 2010 & 2011, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult March 25 2011, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus 13cy C3CP March 25 2011, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult March 26 2010, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus ad C3S6 March 13 2010, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult March 25 2011, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus ad C3X3 March 02 2012, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult March 30 2010, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult C3Y1 March 08 2011, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult March 26 2010, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus ad C51C March 26 2010, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult March 12 2011, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus 9CY C5H8 March 21 2009, Ashdod, Israel. Picture: Amir Ben Dov. Also as 3cy. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult March 17 2010, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus 13CY-16CY C5P4 March 2011, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult March 17 2010, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus 15cy C5P8 March 25 2011, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult March 25 2011, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult C60X March 13 2010, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 25 2013, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult C675 March 26 2013, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 29 2014, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus ad C6R7 March 17 2010, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult March 07 2008, Queen Elizabeth N.P. Uganda. Picture: F.J. Broekhuis.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus ad C7YK March 26 2010, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult March 26 2010, Ashdod, Israel. Picture: Amir Ben Dov.
L. f. fuscus 15cy & 17cy C8A6 April 2011 & March 2013, Ashdod, Israel. Picture:  Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult March 26 2010, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult C8M6 March 13 2010, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult March 21 2009, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult C8V6 March 16-17 2010, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 26 2013, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult C8?8 March 16 2010, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 26 2013, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus 16cy C936 March 26 2010, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 28 2013, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus ad C95L March 26 2010, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 25 2013, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus ad C99W March 16 2010, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 25 2013, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus ad C9PV March 16 2010, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 25 2013, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus ad C9XX March 16 2010, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult March 21 2009, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus ad CCAR March 2010 & 2013, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult March 26 2010, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus 16+cy CCJ9 March 12 2011, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult March 12 2011, Ashdod, Israel. Picture: Amir Ben Dov. With heuglini.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus 18cy CE41 March 16 2010, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult March 13 2010, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus ad CEK1 March 26 2010, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult March 13 2010, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus ad CEUP March 26 2010, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult March 16 2010, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult CHSP March 2010 & 2013, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult March 16 2010, Ashdod, Israel. Picture: Amir Ben Dov.
L. f. fuscus 18cy-21cy CJ18 April 2011, March 2013 & March 2014, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult March 25 2011, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus 13cy CJK2 March 26 2010, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 29 2013, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus 13cy CJP3 March 30 2010, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 26 2013, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult CKN0 March 29 2014, Ashdod, Israel. Picture: Amir Ben Dov.  
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult CKX1 March 12 2011, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 29 2013, Ashdod, Israel. Picture: Amir Ben Dov.
L. f. fuscus adult CMME March 26 2013, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult March 12 2011, Ashdod, Israel. Picture: Amir Ben Dov. With heuglini.
L. f. fuscus adult CNMC March 29 2013, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult March 21 2009, Ashdod, Israel. Picture: Amir Ben Dov.
L. f. fuscus adult CNX7 March 25 2013, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult March 12 2011, Ashdod, Israel. Picture: Amir Ben Dov. With michahellis.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus ad CSAP March 13 2010, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult March 26 2010, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus ad CZ9N March 02 2012, Ashdod, Israel. Picture: Amir Ben Dov. Ring code hard to read. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 30 2010, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus ad CT 140.463 March 30 2012, Ashdod, Israel. Picture: Amir Ben Dov. Hit electricity wires. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 28 2013, Ashdod, Israel. Picture: Amir Ben Dov.
L. f. fuscus 30CY HT-054.651 March 17 2012, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 02 2012, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus ad HT 075.158 March 26 2010, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 28 2013, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult HT 091.362 March 13 2010, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 26 2013, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult HT 095.308 March 21 2009, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 28 2013, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus ad HT 217.259 March 02 2012, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 25 2013, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus 1CY & 8CY C00H August 2006 & March 2013, Tampere Finland & Ashdod Israel. Picture Hannu Koskinen & Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 25 2013, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult C0YE March 17 2010, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 25 2013, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult C09P March 26 2010, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 25 2013, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult C1AA March 26 2010, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 23 2012, Eilat, Israel. Picture: Avi Meir.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult C1UU March 26 2013, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 28 2013, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus 4CY & 9CY C27N October-December 2009 & March 2014, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 23 2012, Eilat, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult C28K March 16 2010, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 28 2013, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult C36L March 08 2011, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 28 2013, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus 6CY & 10CY C3WH May 2009 & March 2013, Finland & Israel. Pictures Hannu Koskinen & Amir Ben Dov.  
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus 4cy & 6cy C58J February 2011 & March 2013, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 28 2013, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult C5RW March 29 2013, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 28 2013, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult C61X March 26 2013, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 28 2013, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult C7EP March 25 2011, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 28 2013, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult CC14 March 26 2010, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 28 2013, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult CE43 March 25 2011, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 28 2013, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult CM20 March 13 2010, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 25 2013, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult CN27 March 08 2011, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 28 2013, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus 6-7cy CU42 March 2010-2011, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 28 2013, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus 10+cy CV01 March 12 2011, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 26 2013, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus 10+cy CV13 March 12 2011, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 26 2013, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult CV21 March 2009 & 2014, Ashdod, Israel. Pictures: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 23 2012, Eilat, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus 5cy C09L March 29 2014, Ashdod, Israel. Picture: Amir Ben Dov.  
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult CKJ.0 March 17 2010, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult March 12 2011, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus 6cy CKL.5 March 22-29 2014, Ashdod, Israel. Picture: Amir Ben Dov.  
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus 6cy CLX.5 March 29 2014, Ashdod, Israel. Picture: Amir Ben Dov.  
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult J0X0 2008, 2009 & 2011, Ashdod, Israel. Pictures: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult March 12 2011, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus ad J20E March 26 2010, Ashdod, Israel. Pictures: Amir Ben Dov. Moulting coverts. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult March 21 2009, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus 5cy J449 March 07 2010, Ashdod, Israel. Pictures: Amir Ben Dov. Bird from N Norway. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 23 2012, Eilat, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult J475 March 08 2011, Ashdod, Israel. Picture: Amir Ben Dov. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult, March 23 2012, Eilat, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus 4cy & 8cy J844 May 2009, March 2013 & March 2014, Finland & Israel. Pictures Hannu Koskinen & Amir Ben Dov. Bird from Norway. 1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult March 21 2009, Ashdod, Israel. Picture: Amir Ben Dov.
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus ad KJ87 March 13 2010, Ashdod, Israel. Picture: Amir Ben Dov.Bird from Karelia, Russia. L. f. fuscus adult ---- March 29 2014, Ashdod. Picture Amir Ben Dov.
L. f. fuscus 2CY-7CY JLJ0 February 2009 - April 2014, Ashdod, Israel. Picture: Amir Ben Dov.  
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult KJN9 March 08 2011, Ashdod, Israel. Picture: Amir Ben Dov.  
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult KJNN March 12 2011, Ashdod, Israel. Picture: Amir Ben Dov. Bird from Russia.  
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult KRAS March 30 2010, Ashdod, Israel. Picture: Amir Ben Dov. Bird from Russia.  
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus ad A170 March 26 2010, Ashdod, Israel. Picture: Amir Ben Dov.  
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult A260 March 25 2011, Ashdod, Israel. Pictures: Amir Ben Dov. Bird from Sweden.  
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus ad M056 March 26 2010, Ashdod, Israel. Picture: Amir Ben Dov.  
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus ad MC93 March 30 2010, Ashdod, Israel. Picture: Amir Ben Dov.  
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus ad MR20 March 29 2013, Ashdod, Israel. Picture: Amir Ben Dov.  
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus ad MR51 March 16 2010, Ashdod, Israel. Picture: Amir Ben Dov.  
1cy fuscus in August, ringed in Finland. (81365 bytes)heuglini / fuscus 5CY U0JD 2010-2014, Ashdod, Israel. Picture: Amir Ben Dov & April 2013 Poland, Picture: Mariusz Dabek.  
1cy fuscus in August, ringed in Finland. (81365 bytes)L. f. fuscus adult U7BJ October 2009 & March 2013, Ashdod, Israel. Picture: Amir Ben Dov.Ringed in Israel.